linh, Vu Ngoc, 1. State Key Laboratory for Conservation and Utilization of Subtropical Agro-Bioresources South China Agricultural University, Guangzhou, 510642; 2. Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant Germplasm, Guangzhou, 510642; 3. Guangdong Province Research Center of Woody Forage Engineering Technology, Guangzhou, 510642; 4. College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou, 510642, Viet Nam